Abstract
In a recent article, “Wayward Modeling: Population Genetics and Natural Selection,” Bruce Glymour claims that population genetics is burdened by serious predictive and explanatory inadequacies and that the theory itself is to blame. Because Glymour overlooks a variety of formal modeling techniques in population genetics, his arguments do not quite undermine a major scientific theory. However, his arguments are extremely valuable as they provide definitive proof that those who would deploy classical population genetics over natural systems must do so with careful attention to interactions between individual population members and environmental causes. Glymour’s arguments have deep implications for causation in classical population genetics