What is the relationship between evolutionary contingency and diversity? The evolutionary contingency thesis emphasizes dependency relations and chance as the hallmarks of evolution. While contingency can be destructive of, for example, the fragile and complex dynamics in an ecosystem, I will mainly focus on the productive or causal aspect of contingency for a particular sort of diversity. There are many sorts of diversities: Gould is most famous for his diversity-to-decimation model, which includes disparate body plans distinguishing different phyla. However, structural (...) diversity construed more broadly spans scales, such as organization in and among cells, structural arrangements and biomechanics on various scales, and even the profile of ancestor-descendent relationships or community structure of interactions within ecosystems. By focusing on stochastic processes in contingent evolution, I argue that contingency causes structural diversity. Specifically, I focus on the plurality of structural types of cells, genetic codes, and phyla diversity as case studies. (shrink)

‘Gouldian arguments’ appeal to the contingency of a scientific domain to establish that domain’s autonomy from some body of theory. For instance, pointing to evolutionary contingency, Stephen Jay Gould suggested that natural selection alone is insufficient to explain life on the macroevolutionary scale. In analysing contingency, philosophers have provided source-independent accounts, understanding how events and processes structure history without attending to the nature of those events and processes. But Gouldian Arguments require source-dependent notions of contingency. An account of contingency is (...) source-dependent when it is indexed to some pattern and some process. Positions like Gould’s do not turn on the mere fact of life’s contingency—that life’s shape could have been different due to its sensitivity to initial conditions, path-dependence or stochasticity. Rather, Gouldian arguments require that the contingency is due to particular kinds of processes: in this case, those which microevolutionary theory cannot account for. This source-dependent perspective clarifies both debates about the nature and importance of contingency, and empirical routes for testing Gould’s thesis. (shrink)

There is an "under-representation problem” in philosophy departments and journals. Empirical data suggest that while we have seen some improvements since the 1990s, the rate of change has slowed down. Some posit that philosophy has disciplinary norms making it uniquely resistant to change (Antony and Cudd 2012; Dotson 2012; Hassoun et al. 2022). In this paper, we present results from an empirical case study of a philosophy department that achieved and maintained male-female gender parity among its faculty as early as (...) 2014. Our analysis extends beyond matters of gender parity because that is only one, albeit important, dimension of inclusion. We build from the case study to reflect on strategies that may catalyze change. (shrink)

Recently, philosophers have sought to determine the nature of individuals relevant to evolution by natural selection or evolutionary individuals. The Evolutionary Contingency Thesis is a claim about evolution that emphasizes the role of contingency or dependency relations and chance-based factors in how evolution unfolds. In this article, I argue that if we take evolutionary contingency seriously, then we should be pluralists about the types of individuals in selection.

This paper argues that individuating practices are produced through iterative processes of community and agent-level interactions. This claim will be demonstrated by using three case studies from biology: The structuring of data categories for data collection tables and models; establishing spatial and temporal threshold markers or limits; and the comparative use of phenomenal characteristics as cues for object identification. By drawing from examples of data classification and comparative analysis in the biological sciences, I offer a view about ‘individuation’ as double-barreled (...) according to the method of co-constitutive conceptual analysis. Specifically, the capacity—i.e., the _ability_ to individuate—is co-constituted by community level choices and agent applications: Individuation’s evidential role is generated, revised, and refined by scientific communities and their members through an iterative process of community and agent-level interactions. (shrink)

George G. Simpson (1902–1984) and Stephen J. Gould (1941–2002) were both engaged with the normative – i.e., social, cultural, political, and even ethical – consequences of their evolutionary theorizing. However, there is a normative point of departure between Simpson and Gould’s work in that regard that has received little attention. Yet, their motivations converge into a larger program of resistance and social protection from misconstrued and illegitimate overreaches of the biological sciences leading up to and after the peak of the (...) modern synthesis. (shrink)

In this dissertation, I explore the reach of the Evolutionary Contingency Thesis—a view that emphasizes the role of dependency relations and chance in evolution. Contingency produces diverse biological entities, processes, and mechanisms. I analyze the implications of evolution’s contingency in three areas. First, I address the problem of evolutionary individuality, which concerns the nature of entities that selection acts on. If we accept Lewontin’s 1970 view that individuals are selected, then what exactly are these individuals? I argue that evolutionary contingency (...) and its diversity of outcomes causes a plurality of individuality types. These types of individuals emerge, evolve, and disappear through evolution. Second, I defend a disunified model of major transitions in evolution. Major transitions mark pivotal turning points in life’s history, such as the transition from unicellularity to multicellularity. Philosophers and biologists aim to unify evolution’s transitions by establishing a property all major events share. I argue that evolutionary contingency’s emphasis on diversity supports a disunified model of transitions. That is, we should expect numerous kinds of major transitions given the diverse sets of outcomes caused by evolutionary contingency. Finally, I extend the contingency analysis to a topic in philosophy of medicine: The nature of cancer. It is common to view cancer as a by-product of our evolution as multicelled creatures. However, Lean and Plutynski argue that cancer is also a direct product of evolution by selection. In both cases, cancer is characterized in terms of how it functions as an object of selection. I argue that the diversity among cancers suggests philosophers should shift their focus to cancers’ structural differences rather than only their functional similarities. (shrink)

While Julian S. Huxley’s role in the Eugenics Society is well known, the ways in which his scientific research program intimately intertwined with his broader social views is sometimes overlooked. This paper analyzes Huxley’s earlier and later research centering Individual (1912) and Modern Synthesis (1942) as two case studies in the context of his larger body of work. There currently exists much exceptional literature on Huxley, which is incorporated and reviewed as much as possible. That literature explores the connection between (...) Huxley’s biological views and social views, but there is more to say about the nature of that connection warranting a return to the details of his research program. Huxley aimed to establish the biologist’s role for engineering human evolution towards sets of ideals conceived by the educated elite. (shrink)