A recent analysis of the genomes of Darwin's finches revealed extensive interspecies allele sharing throughout the history of the radiation and identified a key locus responsible for morphological evolution in this group. The radiation of Darwin's finches on the Galápagos archipelago has long been regarded as an iconic study system for field ecology and evolutionary biology. Coupled with an extensive history of field work, these latest findings affirm the increasing acceptance of introgressive hybridization, or gene flow between species, (...) as a significant contributor to adaptive evolution. Here, we review and discuss these findings in relation to both classical work on Darwin's finches and contemporary work showing similar evolutionary signatures in other biological systems. The continued unification of genomic data with field biology promises to further elucidate the molecular basis of adaptation in Darwin's finches and well beyond. (shrink)

A flurry of technological advances in molecular, cellular and developmental biology during the past decade has provided a clearer understanding of mechanisms underlying phenotypic diversification. Building upon such momentum, a recent paper tackles one of the foremost topics in evolution, that is the origin of species‐specific beak morphology in Darwin's finches.1 Previous work involving both domesticated and wild birds implicated a well‐known signaling pathway (i.e. bone morphogenetic proteins) and one population of progenitor cells in particular (i.e. cranial neural crest), (...) as primary factors for establishing beak size and shape. But these results were limited in their ability to explain fully the morphogenetic bases of patterned outgrowth. So in a quest to identify novel genes whose expression correlated with differences in beak anatomy among Darwin's finches, a DNA microarray approach was undertaken using tissues harvested from the Galápagos Islands. The results are striking and point to a protein called calmodulin, which is a mediator of cellular calcium signaling, as a key determinant of beak length. BioEssays 29: 1–6, 2007. © 2006 Wiley Periodicals, Inc. (shrink)

Adaptation by means of natural selection depends on the ability of populations to maintain variation in heritable traits. According to the Modern Synthesis this variation is sustained by mutations and genetic drift. Epigenetics, evodevo, niche construction and cultural factors have more recently been shown to contribute to heritable variation, however, leading an increasing number of biologists to call for an extended view of speciation and evolution. An additional common feature across the animal kingdom is learning, defined as the ability to (...) change behavior according to novel experiences or skills. Learning constitutes an additional source for phenotypic variation, and change in behavior may induce long lasting shifts in fitness, and hence favor evolutionary novelties. Based on published studies, I demonstrate how learning about food, mate choice and habitats has contributed substantially to speciation in the canonical story of Darwin’s finches on the Galapagos Islands. Learning cannot be reduced to genetics, because it demands decisions, which requires a subject. Evolutionary novelties may hence emerge both from shifts in allelic frequencies and from shifts in learned, subject driven behavior. The existence of two principally different sources of variation also prevents the Modern Synthesis from self-referring explanations. (shrink)

Overview * Part I: Introduction * Philip Appleman, Darwin: On Changing the Mind * Part II: Darwin’s Life * Ernst Mayr, Who Is Darwin? * Part III: Scientific Thought: Just before Darwin * Sir Gavin de Beer, Biology before the Beagle * Thomas Robert Malthus, An Essay on the Principle of Population * William Paley, Natural Theology * Jean Baptiste Pierre Antoine de Monet Lamarck, Zoological Philisophy * Charles Lyell, Principles of Geology * John Herschell, The Study of Natural (...) Philosophy * William Whewell, Astronomy and General Physics Considered with Reference to Natural Theology * Alfred Russel Wallace, On the Tendency of Varieties to Depart Indefinitely from the Original Type * Part IV: Selections from Darwin’s Work * The Voyage of the Beagle * o Chapter I. St. Jago-Cape de Verd Island o Chapter XVII. Galapagos Archipelago * On the Tendency of Species to Form Varieties; and On the Perpetuation of Varieties and Species by Natural Means of Selection * o I. Extract from an unpublished Work on Species, by C. Darwin, Esq.... o II.of Letter from C. Darwin, Esq., to Prof. Asa Gray, Boston, U.S., dated Down, September 5th, 1857 * An Historical Sketch of the Progress of Opinion on the Origin of Species, previously to the : Publication of This Work The Origin of Species * o Introduction o Chapter I. Variation under Domestication o Chapter II. Variation under Nature o Chapter III. Struggle for Existence o Chapter IV. Natural Selection o Chapter VI. Difficulties on Theory o Chapter IX. On the Imperfections of the Geological Record o Chapter XIII. Mutual Affinities of Organic Beings: Morphology: Embryology: Rudimentary Organs o Chapter XIV. Recapitulation and Conclusion * The Descent of Man * o Introduction o Chapter I. The Evidence of the Descent of Man from Some Lower Form o Chapter II. On the Manner of Development of Man from Some Lower Form o Chapter III. Comparison of the Mental Powers of Man and the Lower Animals o Chapter VI. On the Affinities and Genealogy of Man o Chapter VIII. Principles of Sexual Selection o Chapter XIX. Secondary Sexual Characters of Man o Chapter XX. Secondary Sexual Characters of Man-continued o Chapter XXI. General Summary and Conclusion * Part V: Darwin’s Influence on Science * THE VICTORIAN OPPOSITION TO DARWIN * o David L. Hull, Darwin and His Critics o Adam Sedgwick, Objections to Mr. Darwin’s Theory of the Origin of Species o Sir Richard Owen, Darwin on the Origin of Species o Fleeming Jenkin, Review of the Origin of Species * VICTORIAN SUPPORTERS OF DARWIN * o Joseph Dalton Hooker, Flora Tasmaniae o Thomas Henry Huxley, On the Relations of Man to the Lowe Animals o Charles Lyell, Principles of Geology o Alfred Russel Wallace, The Debt of Science to Darwin * DARWIN AND THE SHAPING OF MODERN SCIENCE * o Scientific Method in Evolution o National Academy of Sciences, Evolution and the Nature of Science o Richard Dawkins, Explaining the Very Improbable o Lewis Thomas, On the Uncertainty of Science o Noretta Koetge, Postmodernisms and the Problem of Scientific Literary o Richard Dawkins, Science and Sensibility o The Neo-Darwinian Synthesis o Peter Bowler, The Evolutionary Synthesis o The Human Genealogy o Adam Kuper, The Chosen Primate o Ian Tattersall, Out of Africa Again... and Again? o Stephen Jay Gould, The Human Difference o Punctuated Equilibrium o Stephen Jay Gould, [On Punctuated Equilibrium] o Niles Eldredge, The Great Stasis Debate o Rethinking Taxonomy o Kevin Padian, Darwin’s Views of Classification o David L. Hull, Cladistic Analysis o Kevin Padian and Luis M. Chiappe, Cladistics in Action: The Origin of Birds and Their Flight o Evolution as Observable Fact o James L. Gould and William T. Keeton with Carol Grant Gould, How Natural Selection Operates o Peter r. Grant, Natural Selection and Darwin’s Finches o John A. Endler, Natural Selection in the Wild * Part VI: Darwinian Patterns in Social Thought * COMPETITION AND COOPERATION * o Richard Hofstadter, The Vogue of Spencer o Andrew Carnegie, The Gospel of Wealth o Peter Kropotkin, Mutual Aid o Martin A. Nowak, Robert M. May, and Karl Sigmund, The Arithmetics of Mutual Help * NATURE AND NURTURE * o Edward O. Wilson, Sociobiology: The New Synthesis o Stephen Jay Gould, Biological Potentiality vs. Biological Determination o Barbara Ehrenreich and Janet McIntosh, The New Creationism: Biology under Attack * EVOLUTION AND GENDER * o Elizabeth Cady Stanton, The Woman’s Bible o Nancy Makepeace Tanner, On Becoming Human o Evelleen Richards, Darwin and the Descent of Woman o James Eli Adams, Woman Red in Tooth and Claw * EVOLUTION AND OTHER DISCIPLINES * o Edward O. Wilson, [On Consilience] o Randolph H. Nesse and George C. Williams, Evolution and the Origin of Disease o Steven Pinker, How the Mind Works o Steve Jones, The Set within the Skull * Part VII: Darwinian Influences in Philosophy and Ethics * John Dewey, The Influence of Darwin on Philosophy * Daniel C. Dennett, Darwin’s Dangerous Idea: Natural Selection as an Algorithmic Process * Michael Ruse Darwinian Epistemology * Thomas Henry Huxley, Evolution and Ethics * Julian Huxley, Evolutionary Ethics * Michael Ruse and Edward O. Wilson, The Evolution of Ethics * Frans de Waal, Good Natured: The Origin of Right and Wrong in Humans and Other Animals * Matt Ridley, The Origins of Virtue * Part VIII: Evolutionary Theory and Religious Theory * MAINSTREAM RELIGIOUS SUPPORT FOR EVOLUTION * o Pope John Paul II, Message to the Pontifical Academy of Sciences o Central Conference of American Rabbis, On Creationism in School Textbooks o United Presbyterian Church in the U.S.A., Evolution and Creationsim o The Lutheran World Federation, [Statement on Evolution] o The General Convention of the Episcopal Church, Resolution on Evolutionism and Creationism o Unitariuan Universalist Association, Resolution Opposing "Scientific Creationism" * FUNDAMENTALIST CREATIONISM * o Eugene C. Scott, Antievolution and Creationism in the United States o The Scopes Trial o Thomas McIver, Orthodox Jewish Creationists o Harun Yahya, [Islamic Creationism] o Seami Srila Prabhupada, [A Hare Krishna on Darwinian Evolution] o Institute for Creation Research, Tenets of Creationism o Henry M. Morris, Scientific Creationism o Thomas J. Wheeler, Review of Morris o Richard D. Sjolund and Betty McCollister, Evolution at the Grass Roots o Richard D. Sjolund, [Creationism versus Biotechnology] o Betty McCollister, [The Politics of Creationism] o Molleen Matsumara, What Do Christians Really Believe about Evolution? o National Center for Science Education, Seven Significant Court Decisions Regarding Evolution/Creation Issues * PERSONAL INCREDULITY AND ANTIEVOLUTIONISM * o Richard Dawkins, [The Argument from Personal Incredulity] o Phillip E. Johnson, Darwin on Trial o Eugenie C. Scott, Review of Johnson o Michael Behe, Darwin’s Black Box o Robert Dorit, Review of Behe o Michael Ruse, Darwin’s New Critics on Trial * SCIENTISTS’ OPPOSITION TO CREATIONISM * o American Association for the Advancement of Science, Forced Teaching of Creationist Beliefs in Public School Science Education o American Institute of Biological Sciences, Resolution Oposing Creationism in Science Courses o National Association of Biology Teachers, Statement on Teaching Evolution o National Academy of Sciences, Frequently Asked Questions about Evolution and the Nature of Science * FUNDAMENTALIST CREATIONISM AND THE VALUE OF SATIRE * o Michael Shermer, Genesis Revisted: A Scientific Creation Story o Philip Appleman, Darwin’s Ark * Part IX: Darwin and the Literary Mind * DARWIN’S LITERARY SENSIBILITY * o Charles Darwin, Autobiography o L. Robert Stevens, Darwin’s Humane Reading o George Levine, Darwin and Pain: Why Science Made Shakespeare Nauseating o Gillian Beer, Darwin’s Plots * DARWIN’S INFLUENCE ON LITERATURE * o Lionel Stevenson, Darwin among the Poets o George Levine, Darwin among the Novelists o Joseph Wood Krutch, The Tragic Fallacy o Herbert J. Muller, Modern Tragedy o Philip Appleman, Darwin-Sightings in Recent Literature. (shrink)

Discoverer of the theory of evolution by natural selection, and thus the founder of modern evolutionary biology, Charles Darwin is responsible for one of the most fundamental and far‐reaching contributions to the modern scientific world view. Born in 1809 in Shrewsbury into a wealthy Victorian family, Darwin was educated at the universities of Edinburgh and Cambridge. Though his formal education was of little interest to him ‐ “my time was wasted, as far as the academical studies were concerned” (1969, p. (...) 58) ‐ Cambridge did provide the opportunity to pursue his childhood interest in natural history. Upon graduating in 1831, Darwin joined a charting expedition around South America on board HMS Beagle, which was to last for five years. During this time, he made lengthy expeditions inland, studying the flora, fauna, and geology of South America and the Pacific Islands, and accumulating a massive biological and geological collection. These years were critical for Darwin's intellectual development, as he admitted, sowing the seeds of his doubt in the fixity of individual species. Darwin was impressed by a large number of facts that could not be explained on the assumption of the fixity of species, including the close structural similarities between extinct and living species revealed by the fossil record. Most important of all were the tortoises and finches of the Galapagos Islands: each island was inhabited by very similar, yet distinct varieties ‐ clear evidence for Darwin of common ancestry shaped by adaptation to local conditions. The creatures of the Galapagos, Darwin wrote, bring us close “to that great fact ‐ that mystery of mysteries ‐ the first appearance of new beings on this earth” (1839, p. 466). (shrink)

I first visited the Galápagos Islands in June 1998, and little was as I expected. Rather than craggy barrens covered with scrub, lush foliage beautified many islands. Rather than flourishing coastal habitats, surface water temperatures were well above normal, and throughout the archipelago dead or dying sea lions, sea birds, and marine iguanas littered the shores. All of this was the result of increased rainfall and the disruption of the normal upwelling in waters surrounding the archipelago caused by the 1997–1998 (...) El Niño event. When I visited Galápagos again, in February 1999, the islands were suffering one of the driest periods on record, and terrestrial vegetation had died back; but the cool, nourishing upwelling had resumed, and the marine environment was thriving.A “study in contrasts” aptly describes the two extremes in environmental conditions in the Galápagos Islands in 1998 and 1999 and, as Edward Larson captures beautifully in this book, many aspects of their history. Indeed, a theme that emerges in Larson's historical account of the Galápagos is one of irony. Compare Herman Melville's characterization of the islands as “an archipelago of aridities, without inhabitant, history, or hope of either in all time to come” , with modern biologists' and eco‐tourists' view of the Galápagos as “hallowed ground” and “enchanting”; the view of Galápagos as a place where one can so easily see Creation with the view that is is a place where one can so easily see evolution ; the popular belief that Darwin used Galápagos finches as the basis of his theory of natural selection with the reality that a complete picture of Galápagos finch evolution didn't emerge until it was unveiled in 1947 by David Lack, who renamed “Galápagos finches” “Darwin's finches”; the treatment of Galápagos as a land of plenty from which early sailors, buccaneers, and greedy naturalists removed tens of thousands of tortoises with its current status as a land in which a single tortoise named Lonesome George is an icon for conservation worldwide; and its small stature in terms of geographical size with its monumental stature in terms of impact on the history of biological science.Expanding on the last contrast, Larson offers the following insight from Robert Bowman: “No area on Earth of comparable size has inspired more fundamental changes in Man's perspective of himself and his environment than the Galápagos Islands” . As such, the comprehensive history of the place provided by Larson is well justified. It is also fascinating, and Larson tells the tale well—concisely, occasionally with wit, and always with inclusive documentation. The text is divided into three parts, the first covering the period from the discovery of the New World, including Galápagos in 1535, to Darwin's legendary visit in 1835 and the subsequent publication of On the Origin of Species in 1859; the second, the explosion of interest in the archipelago by American and European naturalists in the late nineteenth and early twentieth centuries; and the third, the remarkable transformation of the Galápagos between World War II, during which it served as a military base for the United States, to the present. Following and cited in the text are sixty‐three pages of notes, a veritable gold mine of references to classic and obscure published material that demonstrate both how thoroughly Larson researched the topic and how persistently this small archipelago has pervaded scientific, governmental, religious, and public thought.This book is more than a historical account of scientific exploration in Galápagos; it is also a dramatic story of the history of Galápagos in human thought. By incorporating thoughts and findings of nearly every famous, infamous, and forgettable player in Galápagos exploration in roughly chronological order and in the context of prevailing social or scientific mores, Larson provides an extraordinary glimpse into human nature. Regarding the evolution of human thought in conceptualizing organic evolution, Larson demonstrates that spiritual concerns or other constraints prevent some individuals from progressing intellectually, others eagerly abandon all tradition to embrace new concepts, and still others find middle ground. As noted poignantly, no less than ironically, by Larson, creation science, not evolution, is taught in Galápagos schools, and this once‐pristine archipelago is now bustling with humans and the predictable problems that accompany us. The history of Galápagos, both in terms of its ultimate fate and its influence on human thought, is not finished. The sequel to Evolution's Workshop may be equally intriguing. (shrink)

Darwin’s theory is often illustrated with depictions of different finch beaks or with lined-up skeletons displaying subtle bone-structure changes throughout evolutionary history. In The Deep Histor...

Complementing the publication of Darwin's notebooks and correspondence, this work provides access to the last remaining unpublished source of Darwin manuscript materials. It is a catalog to and a complete transcription of the marks and annotations he made in the margins of his books. The margin comments throw light on Darwin's immediate reactions to his reading matter; further comments on slips of paper stuck inside the covers of the books reveal more considered evaluation. These comments are also fully transcribed. Annotation (...) copyrighted by Book News, Inc., Portland, OR. (shrink)

Charles Darwin's On the Origin of Species is unquestionably one of the chief landmarks in biology. The Origin (as it is widely known) was literally only an abstract of the manuscript Darwin had originally intended to complete and publish as the formal presentation of his views on evolution. Compared with the Origin, his original long manuscript work on Natural Selection, which is presented here and made available for the first time in printed form, has more abundant examples and illustrations of (...) Darwin's argument, plus an extensive citation of sources. (shrink)

On 27th December 1831, HMS Beagle set out from Plymouth under the command of Captain Robert Fitzroy on a voyage that lasted nearly 5 years. The purpose of the trip was to complete a survey of the southern coasts of South America, and afterwards to circumnavigate the globe. The ship's geologist and naturalist was Charles Darwin. Darwin kept a diary throughout the voyage in which he recorded his daily activities, not only on board the ship but also during the several (...) long journeys that he made on horseback in Patagonia and Chile. His entries tell the story of one of the most important scientific journeys ever made with matchless immediacy and vivid descriptiveness. (shrink)

This transcription of notes made by Charles Darwin during the voyage of H. M. S. Beagle records his observations of the animals and plants that he encountered, and provides a valuable insight into the intellectual development of one of our most influential scientists. Darwin drew on many of these notes for his well known Journal of Researches (1839), but the majority of them have remained unpublished. This volume provides numerous examples of his unimpeachable accuracy in describing the wide range of (...) animals seen in the course of his travels, and of his closely analytical approach towards every one of his observations. Only at the very end of the voyage were his first doubts about the immutability of species expressed consciously, but here are to be found the initial seeds of his theory of evolution, and of the fields of behavioural and ecological study of which he was one of the founding fathers. (shrink)

A companion to Charles Darwin's notebooks, 1836-1844 (Cornell U. Pr., 1987). Because Darwin was in the process of formulating his arguments, entries on a single topic might appear in several series of notes at any number of places in various manuscripts. This concordance gathers these citations toge.

ABSTRACTThe historical process underlying Darwin’s Origin of Species did not play a significant role in the early editions of the book, in spite of the particular inductivist scientific methodology it espoused. Darwin’s masterpiece did not adequately provide his sources or the historical perspective many contemporary critics expected. Later editions yielded the ‘Historical Sketch’ lacking in the earlier editions, but only under critical pressure. Notwithstanding the sources he provided, Darwin presented the Origin as an ‘abstract’ in order to avoid (...) giving sources; a compromise he acknowledged and undertook to set right in later editions, yet failed to provide throughout the six editions under his supervision. Darwin’s reluctance to publish the historical context of his theory and his sources, particularly sources which were also ‘precursors’, may be attributed as much to the matter of intellectual ownership as science, or even good literary practice. Of special concern to Darwin were issues of priority or originality over ‘descent with modification’ and especially over Natural Selection. Many later historians have argued that Darwin was unaware of the work of his precursors on Natural Selection. Darwin’s theory was an example of independent discovery, albeit along with such obscure precursors as Matthew or Wells, who were unknown to Darwin until after the publication of the Origin. Both Matthew and Wells had a medical education, like James Hutton or Erasmus Darwin earlier in the eighteenth century, or even Charles Darwin. Evolutionary theory, at least in Britain was a product largely of the medical evolutionists rather than the natural historians which ‘history’ has chosen to select for the focus of attention; and among the medical evolutionists the figure of John Hunter stands out as theorist, experimentalist and teacher: the medical evolutionists were predominantly the product of Hunter’s legacy or of the medical profession and particularly the Scottish Universities. Much recent Darwin scholarship has focused on the private Notebooks, to establish Darwin’s discovery of Natural Selection around 1837–1838 and demonstrate Darwin’s ignorance of his precursors; requiring an explicit acknowledgement by Darwin as the legitimate substantiation of any claim to prior influence. The precursors have been categorized as uniformly obscure or irrelevant to the science of evolution which may be defined exclusively as ‘Darwinian’. The inclination to acknowledge influences, however was not something Darwin was gratuitously given to doing, especially on matters of priority. The Notebooks are not Darwin’s private thoughts; from an early stage he considered them incipient public documents and later sought to protect them as proof of his originality. William C. Wells was not an obscure thinker, but a celebrated scientist whom Herschel, Darwin’s guide to scientific methodology, had recommended as providing a model of scientific method. Darwin discovered Wells through Herschel, and quickly acquired a copy of Wells’ recommended work, no later than 1831, and held it thereafter in his library at Down House. This book, the 1818 edition of Wells’ Two Essays contains a third essay, Wells’ account of Natural Selection. Later, in the Descent of Man Darwin acknowledged his separate discovery of the correlation of colour and disease immunity in man, also earlier recounted by Wells. (shrink)

Like other aspiring geologists in the 1830s, Darwin focused heavily on the rising and falling of the earth's crust. I use his time in the Andes to underscore the importance he placed on larger questions of vertical movement, which mountains helped to solidify in his mind. His most impressive ramblings occurred in 1835 on two high passes in the Andes. Prior to his upland journey, he was well prepared to see the gradual movement of the earth's crust, but his time (...) in the mountains honed his vertical vision. His actual travels up and over enabled him to think more clearly about rise and fall, and how corresponding geographies could link the past to the present. Mountains were instructive backdrops to his theorizing, partly because he traveled through them, but also because his time there, linked with other geographies, became a means to foster his thinking both spatially and temporally. He did not think about mountains, but with them and through them. He is a good example of the broader vertical consciousness that directed many sciences at this time. (shrink)

The publication of Darwin's On the Origin of Species in 1859 ignited a public storm he neither wanted nor enjoyed. Having offered his book as a contribution to science, Darwin discovered to his dismay that it was received as an affront by many scientists and as a sacrilege by clergy and Christian citizens. To answer the criticism that his theory was a theory only, and a wild one at that, he published two volumes in 1868 to demonstrate that evolution was (...) obvious to anyone who cared to look at a bull in a pasture or a dog on a hearth. In response to those who insisted that species were distinct since creation, Darwin pointed to breeders of pigs and pigeons. In reply to those who protested that human intervention is one thing and natural selection another, he argued, "If organic beings had not possessed an inherent tendency to vary, man could have done nothing." To counter those who scorned his descriptions of species in exotic places, he submitted local evidence of cabbages and cauliflower. Based on a wide array of sources, from ancient pictographs to Polish roosters, from skins and from skeletons, from scientific journals and breeding manuals, Darwin assembled a mass of proof--and a hypothesis about species reversion that risked his reputation anew. The Variation of Animals and Plants under Domestication is a two-volume compilation of his thorough and intensive research and the revolutionary conclusions that resulted. The first portion of his work is dedicated to a meticulous analysis of various aspects of plant and animal life, including an inventory of varieties and their physical and behavioral characteristics, investigation of the impact of a species' surrounding environment and the role that both natural and forced changes in this environment have had. Darwin then turns to a richly detailed discussion of the roles of inheritance and crossing in the development of species. A wealth of illustrations further support and enhance his findings. This fascinating, invaluable, and courageous undertaking eventually formed the foundation for our current understanding of evolution. "In science as in politics the victors tend to write the history books. As a result, the record of the past is edited, intentionally or unintentionally, so that it focuses mainly on the precursors of contemporary orthodoxy. Such a focus may accurately represent the genealogy of modern ideas, but it almost inevitably misrepresents the historical experience of their progenitors... Even the powerful, persuasive, and ultimately triumphant theory of evolution by natural selection required not only defense, but repeated buttressing and revision. Variation showed Darwin hard at work on this rearguard action, using the materials he had at hand... His information was gleaned from the observations of fanciers, breeders, and amateur naturalists, as well as from the treatises of those on the cutting edge of zoology and botany. As hindsight narrows the historical spotlight, it imposes its own sense of hierarchy on the preoccupations of the past. But Darwin was interested in all of these topics, valued all of these sources, and belonged, to a greater or lesser extent, to all of these communities."--from the Introduction. (shrink)

It is not justifiable to accuse Darwin of conscious or unconscious plagiarism. This charge is contrary to the historical evidence and to the extensive information that we have about his character. When Darwin listed the writers on the origin of species by natural selection before himself, he did not mention Blyth, and this omission did not disturb the cordial relations between Darwin and Blyth. Blyth continued to supply Darwin with information which Darwin used in his later publications with due acknowledgment (...) to Blyth. For example, in The Descent of Man, Darwin cited Blyth: “Mr. Blyth, as he informs me, saw Indian crows feeding two or three of their companions which were blind.”63 Blyth felt no resentment. If he did, he would have so informed Darwin. Blyth did not regard himself as in any sense a predecessor of Darwin and he certainly did not resent Darwin as a plagiarizer of himself. Moreover, Darwin went to a great deal of trouble to find his own predecessors and to give them proper credit.64After Darwin had completed his work on natural selection, he wrote a letter to the Reverend Baden Powell in which he clearly showed recognition of the contribution of others to his own work:No educated person, not even the most ignorant, could suppose I mean to arrogate to myself the origination of the doctrine that species had not been independently created. The only novelty in my work is the attempt to explain how species became modified, and to a certain extent how the theory of descent explains certain large classes of facts; and in these respects I received no assistance from my predecessors.65 *** DIRECT SUPPORT *** A8402011 00002. (shrink)

Various claims have been made, recently, that Darwin's argumentation in the Origin instantiates and so supports some general philosophical proposal about scientific theorizing, for example, the "semantic view". But these claims are grounded in various incorrect analyses of that argumentation. A summary is given here of an analysis defended at greater length in several papers by the present author. The historical and philosophical advantages of this analysis are explained briefly. Darwin's argument comprises three distinct evidential cases on behalf of natural (...) selection, cases, that is, for its existence, its adequacy and its responsibility. Theorizing, today, about evolution by natural selection involves a similar structure of evidential and explanatory concerns. (shrink)

The first annotated edition of the book that shocked the Victorian world and continues to generate controversy today.

The philosophical or metaphysical architecture of Darwin's theory of evolution by natural selection is analyzed and diflussed. It is argued that natural selection was for Darwin a paradigmatic case of a natural law of change — an exemplar of what Ghiselin (1969) has called selective retention laws. These selective retention laws lie at the basis of Darwin's revolutionary world view. In this essay special attention is paid to the consequences for Darwin's concept of species of his selective retention laws. Although (...) Darwin himself explicity supported a variety of nominalism, implicit in the theory of natural selection is a solution to the dispute between nominalism and realism. It is argued that, although implicit, this view plays a very important role in Darwin's theory of natural selection as the means for the origin of species. It is in the context of these selective retention laws and their philosophical implications that Darwin's method is appraised in the light of recent criticisms, and the conclusion drawn that he successfully treated some philosophical problems by approaching them through natural history. Following this an outline of natural selection theory is presented in which all these philosophical issues are highlighted. (shrink)

We lay out the fatalist’s argument, making sure to clarify which dialectical moves are available to the libertarian. We then offer a more robust presentation of Ockhamism, responding to obvious objections and teasing out the implications of the view. At this point, we discuss presentism and eternalism in more detail. We then present our argument for the claim that the libertarian cannot take Ockham’s way out of the fatalism argument unless she rejects presentism. Finally, we consider and dispense with objections (...) to our argument. In the end, it ought to be clear that the libertarian must make a choice between Ockham’s way out and presentism. (shrink)

This essay examines Darwin's positiva use of theology in the first edition of the Origin of Species in three steps. First, the essay analyses the Origin's theological language about God's accessibility, honesty, methods of creating, relationship to natural laws and lack of responsibility for natural suffering; the essay contends that Darwin utilized positiva theology in order to help justify descent with modification and to attack special creation. Second, the essay offers critical analysis of this theology, drawing in part on Darwin's (...) mature ruminations to suggest that, from an epistemic point of view, the Origin's positiva theology manifests several internal tensions. Finally, the essay reflects on the relative epistemic importance of positiva theology in the Origin's overall case for evolution. The essay concludes that this theology served as a handmaiden and accomplice to Darwin's science. (shrink)

Charles Robert Darwin (1809–1882) has been widely recognized since his own time as one of the most influential writers in the history of Western thought. His books were widely read by specialists and the general public, and his influence had been extended by almost continuous public debate over the past 150 years. New York University Press's new paperback edition makes it possible to review Darwin's public literary output as a whole, plus his scientific journal articles, his private notebooks, and his (...) correspondence. This is complete edition contains all of Darwin's published books, featuring definitive texts recording original pagination with Darwin's indexes retained. The set also features a general introduction and index, and introductions to each volume. (shrink)

Summary The common view has been that Darwin regarded species as artificial and arbitrary constructions of taxonomists, not as distinct natural units. However, in his transmutation notebooks he clearly subscribed to the reality of species, on the basis of the criterion of non-interbreeding. A consequence of this biological species concept was his identification of the acquisition of reproductive isolation as the mark of the completion of speciation. He developed in the notebooks a theory of geographic speciation on the grounds of (...) (a) an empirical correlation between barriers and the geographical distributions of allied species, (b) the analogy of artificial isolation in the formation of new domesticated varieties, and (c) the logical consequence of blending inheritance. A brief discussion of Darwin's species concept in later writings concludes the paper. (shrink)

We view an individual’s legitimacy judgment as an attitude. It is influenced by a personal belief system composed of global values and domain-specific beliefs, consistent with the value–attitude system in marketing. Our context is the legitimacy of the Canadian oil sands industry. We hypothesize that an individual’s legitimacy judgment may be influenced by three domain-specific beliefs: the credibility of the industry, environmental non-government organizations, and the mass media. We also examine two global values associated with sustainable development: concern for the (...) environment and concern for economic development. These are expected to have negative and positive effects, respectively, on an individual’s legitimacy judgment and on industry credibility. We also examine if these relationships are influenced by proximity to the oil sands. We test our theory using a mixed methods research design and a structural equation analysis on a survey of 853 residents of Edmonton and Toronto, Canada. We find support for most of our hypotheses. (shrink)

A treasure trove of illuminating and entertaining quotations from the legendary naturalist Here is Charles Darwin in his own words—the naturalist, traveler, scientific thinker, and controversial author of On the Origin of Species, the book that shook the Victorian world. Featuring hundreds of quotations carefully selected by world-renowned Darwin biographer Janet Browne, The Quotable Darwin draws from Darwin’s writings, letters to friends and family, autobiographical reminiscences, and private scientific notebooks. It offers a multifaceted portrait that takes readers through his (...) youth, the famous voyage of the Beagle, the development of his thoughts about evolution, his gradual loss of religious faith, and the time spent turning his ideas into a well-articulated theory about the natural origin of all living beings—a theory that dangerously included the origin of humans. The Quotable Darwin also includes many of the key responses to Darwin’s ideas from figures across the social spectrum, scientists and nonscientists alike—and criticism too. We see Darwin as an innovative botanist and geologist, an affectionate husband and father, and a lively correspondent who once told his cousin that he liked to play billiards because “it drives the horrid species out of my head.” This book gives us an intimate look at Darwin at work, at home, as a public figure, and on his travels. Complete with a chronology of Darwin’s life by Browne, The Quotable Darwin provides an engagingly fresh perspective on a remarkable man who was always thinking deeply about the natural world. (shrink)

For nearly five years, from Dec. 27, 1831, until Oct. 2, 1836, I served as naturalist aboard the H.M.S. Beagle, exploring. During that voyage I was much amazed by how the various types of organisms were distributed around South America, and how the animals and plants presently living on that continent are related to those found only as fossils in the geological record elsewhere. These facts, as will be seen in later chapters, seemed to me to throw some light on (...) the origin of species-that "mystery of mysteries," as it has been called by one of our greatest scientists, John Herschel. After I returned home, it occurred to me in 1837 that I might be able to help address this great question by patiently accumulating and reflecting on all sorts of facts that might have any bearing on it. Finally, after five years of work, I allowed myself to speculate on the subject and wrote up some brief notes. I enlarged these in 1844 into a sketch of the conclusions that seemed to be most probable from the evidence I had collected. Over the subsequent 15 years I have steadily pursued the same object: trying to understand how new species come about. I hope you will excuse me for entering these personal details of my work, as I give them only to show that I have not been hasty in coming to a decision. (shrink)

In Darwin’s Sacred Cause, Adrian Desmond and James Moore contend that “Darwin would put his utmost into sexual selection because the subject intrigued him, no doubt, but also for a deeper reason: the theory vindicated his lifelong commitment to human brotherhood”. Without questioning Desmond and Moore’s evidence, I will raise some puzzles for their view. I will show that attention to the structure of Darwin’s arguments in the Descent of Man shows that they are far from straightforward. As (...) Desmond and Moore note, Darwin seems to have intended sexual selection in non-human animals to serve as evidence for sexual selection in humans. However, Darwin’s account of sexual selection in humans was different from the canonical cases that Darwin described at great length. If explaining the origin of human races was the main reason for introducing sexual selection, and if sexual selection was a key piece of Darwin’s anti-slavery arguments, then it is puzzling why Darwin would have spent so much time discussing cases that did not really support his argument for the origin of human races, and it is also puzzling that his argument for the origin of human races would be so poor. (shrink)

Gathers selections from nine of Darwin's most important books, including writings about coral reefs, the Galapagos Islands, evolution, emotions, and flowers.

In On the Origin of Species, Charles Darwin put forward his theory of natural selection. Conventionally, Darwin's argument for this theory has been understood as based on an analogy with artificial selection. But there has been no consensus on how, exactly, this analogical argument is supposed to work – and some suspicion too that analogical arguments on the whole are embarrassingly weak. Drawing on new insights into the history of analogical argumentation from the ancient Greeks onward, as well as on (...) in-depth studies of Darwin's public and private writings, this book offers an original perspective on Darwin's argument, restoring to view the intellectual traditions which Darwin took for granted in arguing as he did. From this perspective come new appreciations not only of Darwin's argument but of the metaphors based on it, the range of wider traditions the argument touched upon, and its legacies for science after the Origin. (shrink)

This paper examines the way Charles Darwin applied his domestic breeding analogy to the practical workings of species evolution: that application, it is argued, centered on Darwin's distinction between methodical and unconscious selection. Methodical selection, which entailed pairing particular individuals for mating purposes, represented conditions of strict geographic isolation, obviously useful for species multiplication (speciation). By contrast, unconscious selection represented an open landmass with a large breeding population. Yet Darwin held that this latter scenario, which often would include multiple ecological (...) subdistricts and thus partial isolation, was better suited for speciation than were isolated conditions. At the same time, many passages in Darwin's writings that apparently portrayed phyletic evolution exclusively (these including references to unconscious selection), actually applied to speciation as well, for phyletic change in a single district could constitute a local manifestation of a larger common-descent pattern. This generic use of "phyletic" change was reflected in Darwin's deployment of the unconscious selection analogy in his published writings as well as in his dispute with Moritz Wagner over the necessity of geographic isolation for speciation. We can thus understand Darwin's otherwise puzzling declaration in The Origin of Species that unconscious selection was 'more important' than the methodical approach. (shrink)

Since the time, about a century ago, when DeMorgan, Boole and Jevons, inaugurated the study of the logic of numerically definite reasoning, no one has been concerned to establish the validity rules for a very general type of numerically definite inference which is a strong analogue of the classical syllogism. The reader will readily agree that the traditional rules of syllogistic inference cannot even begin to decide whether the following proportionally quantified syllogism is a valid argument: at most 4/7 p (...) are at most 2/3 not-m and at least 3/5 s are precisely 7/8 m, hence, at most 1/3 s are p. Likely reasons, of broad methodological interest, why this readily definable type of formal inference has been neglected will presently be proposed. Among the masters of twentieth century logic, Hilbert and Bernays have, to be sure, meticulously shown how such numerically definite propositions as “at least 7 individuals have the property p” can be expressed in the predicate calculus with the use of the identity relation. They do not, since their concern with such propositions is indirect, proceed to establish a systematic calculus of classes whose “size” or extent is numerically definite. As we shall always notice, traditional logic and even ordinary Boolean algebra prefer to generalize about the laws governing the interrelationship of classes in inference systems without making any hypotheses concerning class “size” or extent except by the classical quantifier “some”, which quantifier admits the measure or extent of a class only in terms of the proposition that a given class contains or does not contain at least one member. Such a restricted admission of the relevance for inference of class size is among the fundamental reasons for the neglect of proportional quantifiers, i.e., quantifiers where numerical value is any rational number r such that 0/100 ≤ r ≤ 100/100. It can, of course, also be pointed out that the traditional syllogism and ordinary Boolean algebra do not i.e. measure, numerically define, the degree of inclusion of one class in another. Inclusion is taken either as total or partial, with no discrimination of the degree of partial inclusion. (shrink)

The so-called Mind argument aims at the conclusion that agents act freely only if determinism is true. The soundness of this argument entails the falsity of libertarianism, the two-part thesis that agents act freely, and free action and determinism are incompatible. In this paper, I offer a new formulation of the Mind argument. I argue that it is true by definition that if an agent acts freely, either (i) nothing nomologically grounds an agent’s acting freely, or (ii) the consequence argument (...) for incompatibilism is unsound. I define the notion of nomological grounding, and argue that unless an agent’s acting freely is nomologically grounded, unacceptable consequences follow. I then argue that if agents act freely and the consequence argument is sound, a vicious regress ensues. I conclude by considering the libertarian’s dialectical options. (shrink)

The central role played by Darwin's analogy between selection under domestication and that under nature has been adequately appreciated, but I have indicated how important the domesticated organisms also were to other elements of Darwin's theory of evolution-his recognition of “the constant principle of change,” for instance, of the imperfection of adaptation, and of the extent of variation in nature. The further development of his theory and its presentation to the public likewise hinged on frequent reference to domesticates.We have seen (...) that Darwin's reliance on the analogy between domesticated varieties and wild species was a bold and original step, in light of contemporary views on the nature of domesticates. However, as Darwin undoubtedly foresaw, his reliance on the analogy created difficulties as well as solving problems, and these began with his Malthusian codiscoverer of the principle of natural selection, Alfred Russel Wallace. Wallace's paper “On the Tendency of Varieties to Depart Indefinitely from the Original Type,” presented to the Linnean Scoiety along with the first public unveiling of Darwin's theory, states: We see, then, that no inferences as to varieties in a state of nature can be deduced from the observation of those occurring among domestic animals. The two are so much opposed to each other in every circumstance of their existence, that what applies to the one is almost sure not to apply to the other. Domestic animals are abnormal, irregular, artificial; they are subject to varieties which never occur and never can occur in a state of nature.62Much has been made of the similarity of views of Darwin and Wallace, but this quotation surely reveals how utterly different their views were on what to Darwin was an important matter. Several critics of the Origin saw Darwin's reliance on the domesticates as his Achilles heel. As Young has pointed out, Samuel Wilberforce included the following passage in his attack on the Origin: Nor must we pass over unnoticed the transference of the argument from the domesticated to the untamed animals. Assuming that man as the selector can do much in a limited time, Mr. Darwin argues that Nature, a more powerful, a more continuous power, working over vastly extended ranges of time, can do more. But why should Nature, so uniform and persistent in all her operations, tend in this instance to change? Why should she become a selector of varieties?63Another critic, Fleeming Jenkin, found the analogy a weakness in Darwin's theory because of the limited extent of variation in any one direction in domestic animals and plants.64 We have already seen that Darwin had confided a similar view to his notebook thirty years earlier, but changed his mind as a result of his profound study of domesticates. De Beer's reference to “an English country gentleman's knowledge of domestic plants and animals and their breeding”65 fails totally to recognize the originality and depth of Darwin's knowledge of domesticates.Why did Darwin, against the currents of his time, rely so heavily on mankind's experience with domesticated organisms to shape his theory about species in nature? On reason is that only with domesticates was an approach that came close to experimental verification possible. Darwin fully realized the inadequacies of the experiment, as is emphasized by his repeated contrasting of selection under nature and selection by man. Yet the extensive experience and data of plant and animal breeders offered the only reliable base against which Darwin could continually challenge his views. As he wrote in the introduction to Variation, with domestication, “man ... may be said to have been trying an experiment on a gigantic scale.”66 Given Darwin's high opinion of the quantitative work of Malthus and Quetelet (as emphasized by Schweber),67 and his unremitting efforts to secure data by which to test his theories, it was inevitable that he should attach high significance to domesticated varieties. John Tyndall, in his Belfast address of 1874, said: “The strength of the doctrine of Evolution consists, not in experimental demonstration (for the subject is hardly accessible to this mode of proof), but in its general harmony with scientific thought.”68 Darwin would have agreed with the latter thought, but I think he would have challenged the preceding one on the grounds that long experience with domesticated varieties did provide an element of experimental demonstration. It gave him confidence in his theory, and he used his vast knowledge of artificial selection boldly and creatively. (shrink)

Darwin's book On the Origin of Species by Means of Natural Selection (henceforth The Origin) abounds with metaphors. In fact, the very theory of natural selection is couched in a system of metaphors that exhibit striking consistency and coherence. I argue that the phenomenon for which Darwin tries to detect the basic mechanisms, that is, biological evolution, involves vast, indeterminate, and ambiguous observations that are difficult to subject to the empirical methods. This fact motivates Darwin's extensive use of metaphors to (...) organize his observations, structure the vague concepts, and ultimately render his observations meaningful and intelligible. I demonstrate that Darwin's metaphors, as far as they are elements of Idealized Cognitive Models, prove valuable in achieving this goal. First, I identify the conceptual metaphors underlying the main metaphorical expressions and show how these conceptual metaphors give rise to entailments and inferences central to Darwin's theory. The conceptual metaphors I identify are “NATURE IS A MOTHER,” “NATURE IS A BREEDER,” “LIFE IS WAR,” “LIFE IS A RACE,” and “EVOLUTION IS PROGRESS.” I also comment briefly on the conceptual blending involved in these metaphorical mappings. I then turn to characterizing the idealized cognitive models (ICMs) that function as the pre-conceptual ground in virtue of which Darwin's metaphors are meaningful, coherent, and helpful to scientific thinking. These ICMs are the ICM of STRUGGLE and the image-schema of SOURCE-PATH-GOAL. (shrink)